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palmbeachpop
I'm told that truck drivers using their own fuel try to stick as close to 2200rpm as they can for best diesel efficiency. I suspect gasoline engines do about the same.
Lee_T
A "Car and Driver" editor once went even further - he said that driving in such a high gear that the gas pedal is floored yields the highest mileage because when the pedal is floored is when the engine has the highest volumetric efficiency. This was back in the days of carburetors so when the pedal is floored the throttle plate is completely open. I don't know if this applies to fuel injected engines.
Truck Driver
Big truck max out 2100 rpms - Many have trucks governed down to 1800 rpms. Every engine is a little different but figure around 1300 - 1400 rpm is were big trucks get best fuel.
Oldtimer2002
A gasoline internal combustion engine gets better efficiency when the vacuum pumping losses are reduced by opening the throttle valve wide open. Low rpm gives best fuel economy with a low overlap camshaft. Wide open efficiency with a carbuerator is difficult as the metering orifices usually open up mechanically with the throttle plate movement. Carburetors also have an accelorator pump that squirts more fuel with rapid throttle plate movement.
Wakebe
Sorry, while final answer may be right, didn't care for this explanation, maybe better without the cycling analogy. Still, there is an optimal RPM for efficiency whether its a piston engine, turbo-shaft, or human running or human on a bicycle. This seems to imply low RPM always more efficient, which is obviously not true, other inefficiencies will take over at lower cadence/rpm. In cycling optimal tends to be 90-100 RPM, running about 180 strides/min (with 2 strides per cycle its about equivalent to the 90-100 RPM). Same with car engines, there will be an optimal. Adding more speeds to a bicycle allows more variability to stay at/near optimal RPM depending on the conditions (wind, incline, terrain, etc).
Gearcat
Just because a transmission has six speeds doesn't mean it has higher or lower gear ratios. It's the same at each end with more gears in between. Okay, maybe they are slightly different at top gear, but not that much than a three speed.
CarFan
This is a bit more complicated than the bicycle analogy in that there are other factors involved as well. Torque/ gear ratios, engine size and the car's aerodynamic efficiency ratio. A Corvette can be 'tweaked' to get optimal efficiency driving at higher revs and thus higher highway speeds than a Camry. In addition low revs at high torque won't get you up a hill unless your driving a diesel. (Last time I rode my bike up a hill in a low gear I couldn't make it. because it required more torque/ drive energy, than I could provide.) Overall the idea is to keep the engine 'working' at the correct RPM it has been designed for. So his car may very well get better mileage at 3000 RPM and she might get better mileage at 2500 RPM (Although I think that 3K is a bit high for efficient operation.)
ShiftKicker
@palmbeachpop: Diesels get better fuel economy at 2200 because that is close to their torque peak. By this reasoning, gas engines would be most efficient at 4500 rpm or so. However, by this speed in top gear, wind resistance reduces efficiency so that gas engines would be more efficient at 2000-3000 rpm in the same gear.
ShiftKicker
Unless it's a turbocharged engine.
richalt
I have a car with an MPG readout and I definitely get the best mileage when I feather touch the accelerator and use the minimum amount of pressure. Get that engine down to the lowest, lowest RPM that keeps the car moving in an optimal speed range of 35 to 50 MPH. Slower than 35, then engine is producing more power than needed, even at idle. Faster than 50 the engine requires more accelerator pedal to maintain the speed. Optimum is the speed where you can sustain the MPH with a feather touch on the accelerator.